Fact: current life forms
differ
from those of previous times,
and
are 'descended' from them.
Evolution is descent with
modification.
["Descent" in the genealogical
sense]
Fact: current life
forms are
extremely variable,
both
within and among species.
Evolution accounts for variation
among living
organisms.
[Variation occurs in time
and space]
"Theory of
Evolution"
The explanation
of observed patterns of temporal & spatial variation
in
terms of biological and physical processes.
Natural
Selection provides a mechanism for Evolution:
Modern
evolutionary theory seeks to clarify this mechanism.
The observable
order in Nature is due to common
descent from an ancestor:
Organisms
resemble each other because they are related.
Fact:
All living things are related (the basic fact of
biology):
The
degree
of relationship provides a basis for "natural
classification":
Taxonomy
should reflect the phylogeny of organisms.
I summon the supernatural beings
Who first contrived
The transmorgifications
In the stuff of life.
You did it for your own amusement.
Descend again, be pleased to
reanimante
This revival of those marvels.
Reveal, now, exactly
How they were performed
From the beginning
Up to this moment.
Ovid:
Metamorphoses (trans. Ted Hughes)
The
Classical
Tradition: Plato & Aristotle
Theory
of Forms (essences, eidos)
'real' objects are manifestations of 'ideal' forms
variation
is illusory [see Plato "The
Republic"]
Dichotomy:
the world is composed of paired opposites
"A" versus "not A" classes
good
/ bad, right / wrong, up / down, light / dark, male / female,
etc.
e.g., vertebrates vs. invertebrates
Aristotle
(384-322 BCE) - "Father of Biology"
Five
books on zoology ("Generation
of Animals")
Biological
structures have purpose: Efficient versus Final Causes
Natural
Theology (John Ray
1627-1705)
"The Wisdom of God, Manifested in
His Creation"
(1691)
'Ideal'
forms exist in the Mind of God:
'real'
world created by God (Genesis
1:1)
The
study of nature is a pious activity
Scala Naturae: the "Great
Chain of Being"
Creation
is an infinitely graduated progressive series
Time
scale is short (ca. 6,000 years)
Species
are static: no new forms, no change, no extinction
Linnean
Taxonomy (Carl von Linne
[Carolus
Linnaeus]
1707-1778)
"Systema Naturae"
(1735; 10th
ed. 01 January 1758)
4,162
animals described
binomial nomenclature: genus
+ species names
Worked
"Ad majorem Dei gloriam": for the greater glory of
God
Exploration
creates a Scientific Crisis
New
forms are discovered that don't fit the Scala
Extinctions
have evidently occurred
Variation
is real in space: what about over time?
The Darwinian Revolution
Charles
Darwin
(1809-1882)
BSc
(Cambridge): pre-med
Naturalist
on board HMS "Beagle"
(1831-36)
"The Voyage of the Beagle"
(1839) a
best-seller
Read
Robert Malthus "On
Population" (1838):
population
increases exponentially, resources increase arithmetically
[Married Emma on 29
Jan 1839]
Letter
from Alfred Wallace
(1823-1913) in June 1858
Observation: In any species, more young are born than can possibly survive.
Observation: Yet a species' numbers do not increase without limit.
CONCLUSION: There is a Struggle
for Survival,
and
differential survival &
reproduction occur
within species.
[Darwin: "I use
'struggle' in a large
and metaphorical sense..."].
Observation: Individuals
within
species show variation
that
affects the probability that they will survive this struggle and
leave offspring.
CONCLUSION: Those individuals
that
survive and reproduce do so in consequence
of
their
"adaptively superior"
variation (they
are "more fit")
This
process
of differential survival and reproduction is called Natural
Selection.
Observation: Variation is heritable:
offspring tend to resemble their parents.
[Remember
that Mendelian
genetics was unknown in 1859).
CONCLUSION:
Adaptively
superior variation will be inherited by the offspring
generation.
That
is,
evolution occurs as descent with
modification.
For further reading:
Loren Eisley
(1959).
"Darwin's Century." Doubleday.
William Irvine
(1955).
"Apes, Angels, and Victorians: Darwin, Huxley, & Evolution."
McGraw-Hill.
Ernst Mayr (1994).
"One Long Argument". Harvard University Press.
Gordon Ratray
Taylor
(1963). "The Science of Life." McGraw-Hill.