If two populations have unequal
initial f(A) for
the Red allele, and
exchange individuals at unequal rates m and
m', the expectation is that both populations will
eventually achieve the same allele frequencies by
mixing. However, if the A
allele is subject to differential selection in
the two populations (as in the uni-directional Island
Model), and (or) drift results in random
variation in f(A)
greater than m and m',
a uniform f(A) will
not be reached. The interplay of gene flow, selection, and
drift results in population structure of genetic
variation within species.