"Heads and teeth are drawn to scale, showing the increase in size in the lineages. Front feet and not to scale. Other trends include: a shift from cusps to complex ridges on the grinding surface of the premolars and molars; elongation of the face and of the space between incisors and cheek teeth; an anterior shift of the cheek teeth in the Merychippus-Equus lineage so that they lie forward of the eye; a deeper lower jaw bone; reduction and loss of the side toes; enlargement of the terminal phalanx (hoof); elongation and enlargement of the central metapodial (the longest bone in the foot)." [text from Futuyma 1997; drawing from Stanley 1993, after Simpson 1951]