[page] 22
PART II1.
§§ IV. & V. (ON THE EVIDENCE FROM GEOLOGY.)
I may premise, that according to the view ordinarily received, the myriads of organisms peopling this world have been created by so many distinct acts of creation. As we know nothing of the (illegible) will of a Creator,—we can see no reason why there should exist any relation between the organisms thus created; or again, they might be created according to any scheme. But it would be marvellous if this scheme should be the same as would result from the descent of groups of organisms from [certain] the same parents, according to the circumstances, just attempted to be developed.
With equal probability did old cosmogonists say fossils were created, as we now see them, with a false resemblance to living beings2; what would the Astronomer say to the doctrine that the planets moved (not) according to the law of gravitation, but from the Creator having willed each separate planet to move in its particular orbit? I believe such a proposition (if we remove all prejudices) would be as legitimate as to admit that certain groups of living and extinct organisms, in their distribution, in their structure and in their relations one to another and to external conditions, agreed with the theory
1 In the original ms. the heading is: Part III.; but Part II. is clearly intended; for details see the Introduction. I have not been able to discover where § iv. ends and § v. begins.
2 This passage corresponds roughly to the conclusion of the Origin, see Ed. i. p. 482, vi. p. 661.
GEOLOGY 23
and showed signs of common descent, and yet were created distinct. As long as it was thought impossible that organisms should vary, or should anyhow become adapted to other organisms in a complicated manner, and yet be separated from them by an impassable barrier of sterility1, it was justifiable, even with some appearance in favour of a common descent, to admit distinct creation according to the will of an Omniscient Creator; or, for it is the same thing, to say with Whewell that the beginnings of all things surpass the comprehension of man. In the former sections I have endeavoured to show that such variation or specification is not impossible, nay, in many points of view is absolutely probable. What then is the evidence in favour of it and what the evidence against it. With our imperfect knowledge of past ages [surely there will be some] it would be strange if the imperfection did not create some unfavourable evidence.
Give sketch of the Past,—beginning with facts appearing hostile under present knowledge,—then proceed to geograph. distribution,—order of appearance,—affinities,—morphology &c., &c.
Our theory requires a very gradual introduction of new forms2, and extermination of the old (to which we shall revert). The extermination of old may sometimes be rapid, but never the introduction. In the groups descended from common parent, our theory requires a perfect gradation not differing more than breed(s) of cattle, or potatoes, or cabbages in forms. I do not mean that a graduated series of animals must have existed, intermediate between horse, mouse, tapir3, elephant [or fowl and peacock],
1 A similar passage occurs in the conclusion of the Origin, Ed. i. p. 481. vi. p. 659.
2 See Origin, Ed. i. p. 312, vi. p. 453.
3 See Origin, Ed. i. pp. 280, 281, vi. p. 414. The author uses his experience of pigeons for examples for what he means by intermediate; the instance of the horse and tapir also occurs.
24 GEOLOGY
but that these must have had a common parent, and between horse and this (?) parent &c., &c., but the common parent may possibly have differed more from either than the two do now from each other. Now what evidence of this is there? So perfect gradation in some departments, that some naturalists have thought that in some large divisions, if all existing forms were collected, a near approach to perfect gradation would be made. But such a notion is preposterous with respect to all, but evidently so with mammals. Other naturalists have thought this would be so if all the specimens entombed in the strata were collected1. I conceive there is no probability whatever of this; nevertheless it is certain all the numerous fossil forms fall in(to), as Buckland remarks, not present classes, families and genera, they fall between them: so is it with new discoveries of existing forms. Most ancient fossils, that is most separated (by) space of time, are most apt to fall between the classes-(but organisms from those countries most separated by space also fall between the classes (e.g.) Ornithorhyncus?). As far as geological discoveries (go) they tend towards such gradation2. Illustrate it with net. Toxodon,—tibia and fibula,— dog and otter,—but so utterly improbable is (it), in ex. gr. Pachydermata, to compose series as perfect as cattle, that if, as many geologists seem to
1 The absence of intermediate forms between living organisms (and also as regards fossils) is discussed in the Origin, Ed, i. pp. 279, 280, vi. p. 413. In the above discussion there is no evidence that the author felt this difficulty so strongly as it is expressed in the Origin, Ed. i. p. 299,—as perhaps "the most obvious and gravest objection that can be urged .against my theory." But in a rough summary written on the back of the penultimate page of the ms. he refers to the geological evidence:—"Evidence, as far as it does go, is favourable, exceedingly incomplete,—greatest difficulty on this theory. I am convinced not insuperable." Buckland's remarks are given in the Origin, Ed. i. p. 329, vi. p. 471.
2 That the evidence of geology, as far as it goes, is favourable to the theory of descent is claimed in the Origin, Ed. i. pp. 343-345, vi. pp. 490 -492. For the reference to net in the following sentence, see Note 1, p. 48, of this Essay.
GEOLOGY 25
infer, each separate formation presents even an approach to a consecutive history, my theory must be given up. Even if it were consecutive, it would only collect series of one district in our present state of knowledge; but what probability is there that any one formation during the immense period which has elapsed during each period will generally present a consecutive history. [Compare number living at one period to fossils preserved-look at enormous periods of time.]
Referring only to marine animals, which are obviously most likely to be preserved, they must live where (?) sediment (of a kind favourable for preservation, not sand and pebble)1 is depositing quickly and over large area and must be thickly capped, (illegible) littoral deposits: for otherwise denudation (will destroy them),—they must live in a shallow space which sediment will tend to fill up,—as movement is (in?) progress if soon brought (?) up (?) subject to denudation,—[if] as during subsidence favourable, accords with facts of European deposits2, but subsidence apt to destroy agents which produce sediment3.
I believe safely inferred (that) groups of marine (?) fossils only preserved for future ages where sediment goes on long (and) continuous(ly) and with rapid but not too rapid deposition in (an) area of subsidence. In how few places in any one region like Europe will (?) these contingencies be going on? Hence (?) in
1 See Origin, Ed. i. p. 288, vi. p. 422. "The remains that do become embedded, if in sand and gravel, will, when the beds are upraised, generally be dissolved by the percolation of rain-water."
2 The position of the following is not clear:—" Think of immense differences in nature of European deposits,—without interposing new causes,—think of time required by present slow changes, to cause, on very same area, such diverse deposits, iron-sand, chalk, sand, coral, clay!"
3 The paragraph which ends here is difficult to interpret. In spite of obscurity it is easy to recognize the general resemblance to the discussion on the importance of subsidence given in the Origin, Ed. i. pp. 290 et seq., vi. pp. 422 et seq.
26 GEOLOGY
past ages mere [gaps] pages preserved1. Lyell's doctrine carried to extreme,—we shall understand difficulty if it be asked:—what chance of series of gradation between cattle by (illegible) at age (illegible) as far back as Miocene2? We know then cattle existed. Compare number of living,—immense duration of each period,—fewness of fossils.
This only refers to consecutiveness of history of organisms of each formation.
The foregoing argument will show firstly, that formations are distinct merely from want of fossils (of intermediate beds), and secondly, that each formation is full of gaps, has been advanced to account for fewness of preserved organisms compared to what have lived on the world. The very same argument explains why in older formations the organisms appear to come on and disappear suddenly,—but in [later] tertiary not quite suddenly3, in later tertiary gradually,—becoming rare and disappearing,—some have disappeared within man's time. It is obvious that our theory requires gradual and nearly uniform introduction, possibly more sudden extermination,—subsidence of continent of Australia &c., &c.
Our theory requires that the first form which existed of each of the great divisions would present points intermediate between existing ones, but immensely different. Most geologists believe Silurian* fossils are those which first existed in the whole world,
1 See Note 3, p. 27.
2 Compare Origin, Ed. i. p. 298, vi. p. 437. "We shall, perhaps, best perceive the improbability of our being enabled to connect species by numerous, fine, intermediate, fossil links, by asking ourselves whether,' for instance, geologists at some future period will be able to prove that our different breeds of cattle, sheep, horses, and dogs have descended from a single stock or from several aboriginal stocks."
3 The sudden appearance of groups of allied species in the lowest known fossiliferous strata is discussed in the Origin, Ed. i. p. 306, vi. p. 446. The gradual appearance in the later strata occurs in the Origin, Ed. i. p. 312, vi. p. 453.
4 Compare Origin, Ed. i. p. 307, vi. p. 448.
GEOLOGY 27
not those which have chanced to be the oldest not destroyed,—or the first which existed in profoundly deep seas in progress of conversion from sea to land: if they are first they (? we) give up. Not so Hutton or Lyell: if first reptile1 of Red Sandstone (?) really was first which existed: if Pachyderm2 of Paris was first which existed: fish of Devonian: dragon fly of Lias: for we cannot suppose them the progenitors: they agree too closely with existing divisions. But geologists consider Europe as (?) a passage from sea to island (?) to continent (except Wealden, see Lyell). These animals therefore, I consider then mere introduction (?) from continents long since submerged.
Finally, if views of some geologists be correct, my theory must be given up. [Lyell's views, as far as they go, are in favour, but they go so little in favour, and so much more is required, that it may (be) viewed as objection.] If geology present us with mere pages in chapters, towards end of (a) history, formed by tearing out bundles of leaves, and each page illustrating merely a small portion of the organisms of that time, the facts accord perfectly with my theory3.
1 I have interpreted as Sandstone a scrawl which I first read as Sea; I have done so at the suggestion of Professor Judd, who points out that "footprints in the red sandstone were known at that time, and geologists were not then particular to distinguish between Amphibians and Reptiles."
2 This refers to Cuvier's discovery of Palootherium &c. at Montmartre.
3 This simile is more fully given in the Origin, Ed. i. p. 310, vi. p. 452. "For my part, following out Lyell's metaphor,; I look at the natural geological record, as a history of the world imperfectly kept, and written in a changing dialect; of this history we possess the last volume alone, relating only to two or three countries. Of this volume, only here and there a short chapter has been preserved; and of each page, only here and there a few lines. Each word of the slowly-changing language, in which the history is supposed to be written, being more or less different in the interrupted succession of chapters, may represent the apparently abruptly changed forms of life, entombed in our consecutive, but widely separated formations." Professor Judd has been good enough to point out to me, that Darwin's metaphor is founded on the comparison of geology to history in Ch. i. of the Principles of Geology, Ed. i. 1830, vol. i. pp. 1-4. Professor Judd has also called my attention to another passage,—Principles, Ed. i. 1833, vol. iii. p. 33, when Lyell imagines an historian examining "two buried cities at the foot of Vesuvius, immediately superimposed upon each
28 EXTERMINATION
Extermination. We have seen that in later periods the organisms have disappeared by degrees and [perhaps] probably by degrees in earlier, and I have said our theory requires it. As many naturalists seem to think extermination a most mysterious circumstance1 and call in astonishing agencies it is well to recall what we have shown concerning the struggle of nature. An exterminating agency is at work with every organism: we scarcely see it: if robins would increase to thousands in ten years how severe must the process be. How imperceptible a small increase: fossils become rare: possibly sudden extermination as Australia, but as present means very slow and many means of escape, I shall doubt very sudden exterminations. Who can explain why some species abound more,—why does marsh titmouse, or ring-ouzel, now little change,—why is one sea-slug rare and another common on our coasts,—why one species of Rhinoceros more than another,—why is (illegible) tiger of India so rare? Curious and general sources of error, the place of an organism is instantly filled up.
We know state of earth has changed, and as earthquakes and tides go on, the state must change,— many geologists believe a slow gradual cooling. Now let us see in accordance with principles of [variation] specification explained in Sect. ii. how species would probably be introduced and how such results accord with what is known.
other." The historian would discover that the inhabitants of the lower town were Greeks while those of the upper one were Italians. But he would be wrong in supposing that there had been a sudden change from the Greek to the Italian language in Campania. I think it is clear that Darwin's metaphor is partly taken from this passage. See for instance (in the above passage from the Origin) such phrases as "history...written in a changing dialect"-"apparently abruptly changed forms of life." The passage within [ ] in the above paragraph:—"Lyell's views as far as they go &c.," no doubt refers, as Professor Judd points out, to Lyell not going so far as Darwin on the question of the imperfection of the geological record.
1 On rarity and extinction see Origin, Ed. i. pp. 109, 319, vi. pp. 133,461.
GEOGRAPHICAL DISTRIBUTION 29
The first fact geology proclaims is immense number of extinct forms, and new appearances. Tertiary strata leads to belief, that forms gradually become rare and disappear and are gradually supplied by others. We see some forms now becoming rare and disappearing, we know of no sudden creation: in older periods the forms appear to come in suddenly, scene shifts: but even here Devonian, Permian &c. [keep on supplying new links in chain]-Genera and higher forms come on and disappear, in same way leaving a species on one or more stages below that in which the form abounded.
(GEOGRAPHICAL DISTRIBUTION.)
§ vi. Let us consider the absolute state of distribution of organisms of earth's face.
Referring chiefly, but not exclusively (from difficulty of transport, fewness, and the distinct characteristics of groups) to Mammalia; and first considering the three or four main [regions] divisions; North America, Europe, Asia, including greater part of E. Indian Archipelago and Africa are intimately allied. Africa most distinct, especially most southern parts. And the Arctic regions, which unite N. America, Asia and Europe, only separated (if we travel one way by Behring's St.) by a narrow strait, is most intimately allied, indeed forms but one restricted group. Next comes S. America,—then Australia, Madagascar (and some small islands which stand very remote from the land). Looking at these main divisions separately, the organisms vary according to changes in condition1 of different parts. But besides this, barriers of every kind seem to separate
1 In the Origin, Ed. i. p. 346, vi. p. 493, the author begins his discussion on geographical distribution by minimising the effect of physical conditions. He lays great stress on the effect of barriers, as in the present Essay.
30 GEOGRAPHICAL DISTRIBUTION
regions in a greater degree than proportionally to the difference of climates on each side. Thus great chains of mountains, spaces of sea between islands and continents, even great rivers and deserts. In fact the amount (of) difference in the organisms bears a certain, but not invariable relation to the amount of physical difficulties to transit1.
There are some curious exceptions, namely, similarity of fauna of mountains of Europe and N. America and Lapland. Other cases just (the) reverse, mountains of eastern S. America, Altai (?), S. India (?)2: mountain summits of islands often eminently peculiar. Fauna generally of some islands, even when close, very dissimilar, in others very similar. [I am here led to observe one or more centres of creation3.]
The simple geologist can explain many of the foregoing cases of distribution. Subsidence of a continent in which free means of dispersal, would drive the lowland plants up to the mountains, now converted into islands, and the semi-alpine plants would take place of alpine, and alpine be destroyed, if mountains originally were not of great height. So we may see, during gradual changes4 of climate on a continent, the propagation of species would vary and adapt themselves to small changes
1 Note in the original, "Would it be more striking if we took animals, take Rhinoceros, and study their habitats?"
2 Note by Mr A. R. Wallace. "The want of similarity referred to, is, between the mountains of Brazil and Guiana and those of the Andes. Also those of the Indian peninsula as compared with the Himalayas. In both cases there is continuous intervening land.
"The islands referred to were, no doubt, the Galapagos for dissimilarity from 8. America; our own Islands as compared with Europe, and perhaps Java, for similarity with continental Asia."
3 The arguments against multiple centres of creation are given in the Origin, Ed. i. p. 352, vi. p. 499.
4 In the Origin, Ed. i. p. 366, vi. p. 516, the author does not give his views on the distribution of alpine plants as original but refers to Edward Forbes' work (Geolog. Survey Memoirs, 1846). In his autobiography, Darwin refers to this. "I was forestalled" he says, "in only one important point, which my vanity has always made me regret." (Life and Letters, i. p. 88.)
ALPINE PLANTS 31
causing much extermination1. The mountains of Europe were quite lately covered with ice, and the lowlands probably partaking of the Arctic climate and Fauna. Then as climate changed, arctic fauna would take place of ice, and an inundation of plants from different temperate countries (would) seize the lowlands, leaving islands of arctic forms. But if this had happened on an island, whence could the new forms have come,—here the geologist calls in creationists. If island formed, the geologist will suggest (that) many of the forms might have been borne from nearest land, but if peculiar, he calls in creationist,— as such island rises in height &c., he still more calls in creation. The creationist tells one, on a (illegible) spot the American spirit of creation makes Orpheus and Tyrannus and American doves, and in accordance with past and extinct forms, but no persistent relation between areas and distribution, Geologico-Geograph.—Distribution.
1
(The following is written on the back of a page of the ms.) Discuss one
or more centres of creation: allude strongly to facilities of dispersal
and amount of geological change: allude to mountain-summits afterwards
to be referred to. The distribution varies, as everyone knows,
according to adaptation, explain going from N. to 8. how we
come to fresh groups of species in the same general region, but besides
this we find difference, according to greatness of barriers, in greater
proportion than can be well accounted for by adaptation. (On
representive species see Origin, Ed. i. p. 349, vi. p. 496.)
This very striking when we think of cattle of Pampas, plants (?)
&c. &c. Then go into discussion; this holds with 3 or 4 main
divisions as well as the endless minor ones in each of these 4 great
ones: in these I chiefly refer to mammalia &c. &c. The
similarity of type, but not in species, in same continent has been much
less insisted on than the dissimilarity of different great regions
generically: it is more striking.
(I have here omitted an incomprehensible sentence.) Galapagos Islands,
Tristan d'Acunha, volcanic islands covered with craters we
know lately did not support any organisms. How unlike these islands in
nature to neighbouring lands. These facts perhaps more striking than
almost any others. [Geology apt to affect geography therefore we ought
to expect to find the above.] Geological-geographical distribution. In
looking to past times we find Australia equally distinct. S. America
was distinct, though with more forms in common. N. America its nearest
neighbour more in common, -in some respects more, in some less allied
to Europe. Europe we find (?) equally European. For Europe is now part
of Asia though not (illegible). Africa unknown,—examples, Elephant,
Rhinoceros, Hippopotamus, Hyaena. As geology destroys geography we
cannot be surprised in going far back we find Marsupials and Edentata
in Europe: but geology destroys geography.
32 GEOGRAPHICAL DISTRIBUTION
Now according to analogy of domesticated animals let us see what would result. Let us take case of farmer on Pampas, where everything approaches nearer to state of nature. He works on organisms having strong tendency to vary: and he knows (that the) only way to make a distinct breed is to select and separate. It would be useless to separate the best bulls and pair with best cows if their offspring run loose and bred with the other herds, and tendency to reversion not counteracted; he would endeavour therefore to get his cows on islands and then commence his work of selection. If several farmers in different rinçons1 were to set to work, especially if with different objects, several breeds would soon be produced. So would it be with horticulturist and so history of every plant shows; the number of varieties2 increase in proportion to care bestowed on their selection and, with crossing plants, separation. Now, according to this analogy, change of external conditions, and isolation either by chance landing (of) a form on an island, or subsidence dividing a continent, or great chain of mountains, and the number of individuals not being numerous will best favour variation and selection3. No doubt change could be effected in same country without any barrier by long continued selection on one species: even in case of a plant not capable of crossing would easier get possession and solely
1 Rincon in Spanish means a nook or corner, it is here probably used to mean a small farm.
2 The following is written across the page: "No one would expect a set of similar varieties to be produced in the different countries, so species different."
3 (The following passage seems to have been meant to follow here.) The parent of an organism, we may generally suppose to be in less favourable condition than the selected offspring and therefore generally in fewer numbers. (This is not borne out by horticulture, mere hypothesis; as an organism in favourable conditions might by selection be adapted to still more favourable conditions.)
Barrier would further act in preventing species formed in one part migrating to another part.
ISOLATION 33
occupy an island1. Now we can at once see that (if) two parts of a continent isolated, new species thus generated in them, would have closest affinities, like cattle in counties of England: if barrier afterwards destroyed one species might destroy the other or both keep their ground. So if island formed near continent, let it be ever so different, that continent would supply inhabitants, and new species (like the old) would be allied with that continent. An island generally very different soil and climate, and number and order of inhabitants supplied by chance, no point so favourable for generation of new species2,— especially the mountains, hence, so it is. As isolated mountains formed in a plain country (if such happens) is an island. As other islands formed, the old species would spread and thus extend and the fauna of distant island might ultimately meet and a continent formed between them. No one doubts continents formed by repeated elevations and depressions3. In looking backwards, but not so far that all geographical boundaries are destroyed, we can thus at once see why existing forms are related to the extinct in the same manner as existing ones are in some part of existing continent. By chance we might even have one or two absolute parent fossils.
The detection of transitional forms would be rendered more difficult on rising point of land.
The distribution therefore in the above enumer-
1 (The following notes occur on the back of the page.) Number of species not related to capabilities of the country: furthermore not always those best adapted, perhaps explained by creationists by changes and progress. (See p. 34, note 1.)
Although creationists can, by help of geology, explain much, how can he explain the marked relation of past and present in same area, the varying relation in other cases, between past and present, the relation of different parts of same great area. If island, to adjoining continent, if quite different, on mountain summits,—the number of individuals not being related to capabilities, or how &c.—our theory, I believe, can throw much light and all facts accord.
1 See Origin, Ed. i. p. 390, vi. p. 543. On oscillation see Origin, Ed. i. p. 291, vi. p. 426.
34 GEOGRAPHICAL DISTRIBUTION
ated points, even the trivial ones, which on any other (theory?) can be viewed as so many ultimate facts, all follow (in) a simple manner on the theory of the occurrence of species by (illegible) and being adapted by selection to (illegible), conjoined with their power of dispersal, and the steady geographico-geological changes which are now in progress and which undoubtedly have taken place. Ought to state the opinion of the immutability of species and the creation by so many separate acts of will of the Creator1.
1 (From the back of ms.) Effect of climate on stationary island and on continent, but continent once island. Moreover repeated oscillations fresh diffusion when non-united, then isolation, when rising again immigration prevented, new habitats formed, new species, when united free immigration, hence uniform characters. Hence more forms (on?) the island. Mountain summits. Why not true species. First let us recall in Part I, conditions of variation: change of conditions during several generations, and if frequently altered so much better [perhaps excess of food]. Secondly, continued selection [while in wild state]. Thirdly, isolation in all or nearly all,—as well to recall advantages of.
[In continent, if we look to terrestrial animal, long continued change might go on, which would only cause change in numerical number (? proportions): if continued long enough might ultimately affect all, though to most continents (there is) chance of immigration. Some few of whole body of species must be long affected and entire selection working same way. But here isolation absent, without barrier, cut off such (illegible). We can see advantage of isolation. But let us take case of island thrown up by volcanic agency at some distances, here we should have occasional visitants, only in few numbers and exposed to new conditions and (illegible) more important,—a quite new grouping of organic beings, which would open out new sources of subsistence, or (would) control (?) old ones. The number would be few, can old have the very best opportunity. (The conquest of the indigenes by introduced organisms shows that the indigenes were not perfectly adapted, see Origin, Bd. i. p. 390.) Moreover as the island continued changing,—continued slow changes, river, marshes, lakes, mountains &c. &c., new races as successively formed and a fresh occasional visitant.
If island formed continent, some species would emerge and immigrate. Everyone admits continents. We can see why Galapagos and C. Verde differ (see Origin, Bd. i. p. 398)], depressed and raised. We can see from this repeated action and the time required for a continent, why many more forms than in New Zealand (see Origin, Ed. i. p. 389 for a comparison between New Zealand and the Cape) no mammals or other classes (see however, Origin, Bd. i. p. 393 for the case of the frog). We can at once see how it comes when there has been an old channel of migration,—Cordilleras; we can see why Indian Asiatic Flora,—[why species] having a wide range gives better chance of some arriving at new points and being selected, and adapted to new ends. I need hardly remark no necessity for change.
AFFINITIES AND CLASSIFICATION 35
§ VII. (AFFINITIES AND CLASSIFICATION.)
Looking now to the affinities of organisms, without relation to their distribution, and taking all fossil and recent, we see the degrees of relationship are of different degrees and arbitrary,—sub-genera, -genera,—sub-families, families, orders and classes and kingdoms. The kind of classification which everyone feels is most correct is called the natural system, but no can define this. If we say with Whewell (that we have an) undefined instinct of the importance of organs1, we have no means in lower
Finally, as continent (most extinction (?) during formation of continent) is formed after repeated elevation and depression, and interchange of species we might foretell much extinction, and that the survivor would belong to same type, as the extinct, in same manner as different part of same continent, which were once separated by space as they are by time (see Origin, Ed. i. pp. 339 and 349).
As all mammals have descended from one stock, we ought to expect that every continent has been at some time connected, hence obliteration of present ranges. I do not mean that the fossil mammifers found in S. America are the lineal successors (ancestors) of the present forms of 8. America: for it is highly improbable that more than one or two cases (who will say how many races after Plata bones) should be found. I believe this from numbers, who have lived,—mere (?) chance of fewness. Moreover in every case from very existence of genera and species only few at one time will leave progeny, under form of new species, to distant ages; and the more distant the ages the fewer the progenitors. An observation may be here appended, bad chance of preservation on rising island, the nurseries of new species, appeal to experience (see Origin, Ed. i. p. 292). This observation may be extended, that in all cases, subsiding land must be, in early stages, less favourable to formation of new species; but it will isolate them, and then if land recommences rising how favourable. As preoccupation is bar to diffusion to species, so would it be to a selected variety. But it would not be if that variety was better fitted to some not fully occupied station; so during elevation or the formation of new stations, is scene for new species. But during elevation not favourable to preservation of fossil (except in caverns (?)); when subsidence highly favourable in early stages to preservation of fossils; when subsidence, less sediment. So that our strata, as general rule will be the tomb of old species (not undergoing any change) when rising land the nursery. But if there be vestige will generally be preserved to future ages, the new ones will not be entombed till fresh subsidence supervenes. In this long gap we shall have no record: so that wonderful if we should get transitional forms. I do not mean every stage, for we cannot expect that, as before shown, until geologists will be prepared to say that although under unnaturally favourable condition we can trace in future ages short-horn and Herefordshire (see note 2, p. 26). . After "organs" is inserted, apparently as an afterthought:—" no, and instance metamorphosis, afterwards explicable."
36 AFFINITIES AND CLASSIFICATION
animals of saying which is most important, and yet everyone feels that some one system alone deserves to be called natural. The true relationship of organisms is brought before one by considering relations of analogy, an otter-like animal amongst mammalia and an otter amongst marsupials. In such cases external resemblance and habit of life and the final end of whole organization very strong, yet no relation1. Naturalists cannot avoid these terms of relation and affinity though they use them metaphorically. If used in simple earnestness the natural system ought to be a genealogical (one); and our knowledge of the points which are most easily affected in transmission are those which we least value in considering the natural system, and practically when we find they do vary we regard them of less value2. In classifying varieties the same language is used and the same kind of division: here also (in pine-apple)3 we talk of the natural classification, overlooking similarity of the fruits, because whole plant differs. The origin of sub-genera, genera, &c., &c., is not difficult on notion of genealogical succession, and accords with what we know of similar gradations of affinity in domesticated organisms. In the same region the organic beings are (illegible) related to each other and the external conditions in many physical respects are allied4 and their differences of same kind, and therefore when a new species has been selected and has obtained a place in the economy of nature, we
1 For analogical resemblances see Origin, Ed. i. p. 427, vi. p. 582.
2 "Practically when naturalists are at work, they do not trouble themselves about the physiological value of the characters....If they find a character nearly uniform,...they use it as one of high value," Origin, Ed. i. p. 417, vi. p. 573.
3 "We are cautioned...not to class two varieties of the pine-apple together, merely because their fruit, though the most important part, happens to be nearly identical," Origin, Ed. i. p. 423, vi. p. 579.
4 The whole of this passage is obscure, but the text is quite clear, except for one illegible word.
AFFINITIES AND CLASSIFICATION 37
may suppose that generally it will tend to extend its range during geographical changes, and thus, becoming isolated and exposed to new conditions, will slightly alter and its structure by selection become slightly remodified, thus we should get species of a sub-genus and genus,—as varieties of merino-sheep,—varieties of British and Indian cattle. Fresh species might go on forming and others become extinct and all might become extinct, and then we should have (an) extinct genus; a case formerly mentioned, of which numerous cases occur in Palaeontology. But more often the same advantages which caused the new species to spread and become modified into several species would favour some of the species being preserved: and if two of the species, considerably different, each gave rise to group of new species, you would have two genera; the same thing will go on. We may look at case in other way, looking to future. According to mere chance every existing species may generate another, but if any species, A, in changing gets an advantage and that advantage (whatever it may be, intellect, &c., &c., or some particular structure or constitution) is inherited1, A will be the progenitor of several genera or even families in the hard struggle of nature. A will go on beating out other forms, it might come that A would people earth,—we may now not have one descendant on our globe of the one or several original creations2. External conditions air, earth, water being same3 on globe, and the communication not being perfect, organisms of widely different descent might become adapted to
1 (The exact position of the following passage is uncertain:) "just as it is not likely every present breed of fancy birds and cattle will propagate, only some of the best."
2 This suggests that the author was not far from the principle of divergence on which he afterwards laid so much stress. See Origin, Ed. i. P- HI, vi. p. 134, also Life and Letters, i. p. 84.
3 That is to say the same conditions occurring in different parts of the globe.
38 UNITY OF TYPE
the same end and then we should have cases of analogy1, [they might even tend to become numerically representative]. From this often happening each of the great divisions of nature would have their representative eminently adapted to earth, to (air)2, to water, and to these in (illegible) and then these great divisions would show numerical relations in their classification.
§ VIII. UNITY [OR SIMILARITY] OF TYPE IN THE
GREAT CLASSES.
Nothing more wonderful in Nat. Hist, than looking at the vast number of organisms, recent and fossil, exposed to the most diverse conditions, living in the most distant climes, and at immensely remote periods, fitted to wholely different ends, yet to find large groups united by a similar type of structure. When we for instance see bat, horse, porpoise-fin, hand, all built on same structure3, having bones4 with same name, we see there is some deep bond of union between them5, to illustrate this is the foundation and objects (?) (of) what is called the Natural System; and which is foundation of distinction (?) of true and adaptive characters6. Now this wonderful fact of hand, hoof, wing, paddle and claw being the same, is at once explicable on the principle of some parent-forms, which might either be (illegible) or walking animals, becoming through infinite number of small
1 The position of the following is uncertain, "greyhound and racehorse have an analogy to each other." The same comparison occurs in the Origin, Ed. i. p. 427, vi. p. 583.
2 Air is evidently intended; in the MS. water is written twice.
3 Written between the lines occurs:—" extend to birds and other classes."
4 Written between the lines occurs:—"many bones merely represented."
5 In the Origin, Ed. i. p. 434, vi. p. 595, the term morphology is taken as including unity of type. The paddle of the porpoise and the wing of the bat are there used as instances of morphological resemblance.
6 The sentence is difficult to decipher.
MORPHOLOGY 39
selections adapted to various conditions. We know that proportion, size, shape of bones and their accompanying soft parts vary, and hence constant selection would alter, to almost any purpose (?) the framework of an organism, but yet would leave a general, even closest similarity in it.
[We know the number of similar parts, as vertebrae and ribs can vary, hence this also we might expect.] Also (if) the changes carried on to a certain point, doubtless type will be lost, and this is case with Plesiosaurus1. The unity of type in past and present ages of certain great divisions thus undoubtedly receives the simplest explanation.
There is another class of allied and almost identical facts, admitted by the soberest physiologists, [from the study of a certain set of organs in a group of organisms] and refers (? referring) to a unity of type of different organs in the same individual, denominated the science of "Morphology." The (? this) discovered by beautiful and regular series, and in the case of plants from monstrous changes, that certain organs in an individual are other organs metamorphosed. Thus every botanist considers petals, nectaries, stamens, pistils, germen as metamorphosed leaf. They thus explain, in the most lucid manner, the position and number of all parts of the flower, and the curious conversion under cultivation of one part into another. The complicated double set of jaws and palpi of crustaceans2, and all insects are considered as metamorphosed (limbs) and to see the series is to admit this phraseology. The skulls of the vertebrates are undoubtedly composed of three metamorphosed vertebrae; thus we can understand the strange form of
l In the Origin, Ed. i. p. 436, vi. p. 598, the author speaks of the general pattern "being obscured in the paddles of "extinct gigantic sea-lizards."
2 See Origin, Ed. i. p. 437, vi. p. 599.
40 MORPHOLOGY AND
the separate bones which compose the casket holding man's brain. These1 facts differ but slightly from those of last section, if with wing, paddle, hand and hoof, some common structure was yet visible, or could be made out by a series of occasional monstrous conversions, and if traces could be discovered of (the) whole having once existed as walking or swimming instruments, these organs would be said to be metamorphosed, as it is they are only said to exhibit a common type.
This distinction is not drawn by physiologists, and is only implied by some by their general manner of writing. These facts, though affecting every organic being on the face of the globe, which has existed, or does exist, can only be viewed by the Creationist as ultimate and inexplicable facts. But this unity of type through the individuals of a group, and this metamorphosis of the same organ into other organs, adapted to diverse use, necessarily follows on the theory of descent2. For let us take case of Vertebrata, which if3 they descended from one parent and by this theory all the Vertebrata have been altered by slow degrees, such as we see in domestic animals. We know that proportions alter, and even that occasionally numbers of vertebrae alter, that parts become soldered, that parts are lost, as tail and toes, but we know (that?) here we can see that possibly a walking organ might (?) be converted into swimming or into a gliding organ and so on to a flying organ. But such gradual changes would not alter the unity of type in their descendants, as parts lost and soldered and vertebrae.
1 The following passage seems to have been meant to precede the sentence beginning "These facts":—"It is evident, that when in each individual species, organs are metamorph, a unity of type extends."
2 This is, I believe, the first place in which the author uses the words "theory of descent."
3 The sentence should probably run, "Let us take the case of the vertebrata: if we assume them to be descended from one parent, then by this theory they have been altered &c."
THEORY OF DESCENT 41
But we can see that if this carried to extreme, unity lost,—Plesiosaurus. Here we have seen the same organ is formed (?) (for) different purposes (ten words illegible): and if, in several orders of vertebrata, we could trace origin (of) spinous processes and monstrosities &c. we should say, instead of there existing a unity of type, morphology1, as we do when we trace the head as being the vertebrae metamorphosed. Be it observed that Naturalists, as they use terms of affinity without attaching real meaning, here also they are obliged to use metamorphosis, without meaning that any parent of crustacean was really an animal with as many legs as crustacean has jaws. The theory of descent at once explains these wonderful facts.
Now few of the physiologists who use this language really suppose that the parent of insect with the metamorphosed jaw, was an insect with [more] so many legs, or that the parent of flowering plants, originally had no stamens, or pistils or petals, but some other means of propagation,—and so in other cases. Now according to our theory during the infinite number of changes, we might expect that an organ used for a purpose might be used for a different one by his descendant, as must have been the case by our theory with the bat, porpoise, horse, &c., which are descended from one parent. And if it so chanced that traces of the former use and structure of the part should be retained, which is manifestly possible if not probable, then we should have the organs, on which morphology is founded and which instead of being metaphorical becomes plain and (and instead of being) utterly unintelligible becomes simple matter of fact2.
1 That is "we should call it a morphological fact."
2 In the Origin, Ed. i. p. 438, vi. p. 602, the author, referring to the expressions used by naturalists in regard to morphology and metamorphosis, says "On my view these terms may be used literally."
42 EMBRYOLOGY
(Embryology.) This general unity of type in great groups of organisms (including of course these morphological cases) displays itself in a most striking manner in the stages through which the fotus passes1. In early stage, the wing of bat, hoof, hand, paddle are not to be distinguished. At a still earlier (stage) there is no difference between fish, bird, &c. &c. and mammal. It is not that they cannot be distinguished, but the arteries2 (illegible). It is not true that one passes through the form of a lower group, though no doubt fish more nearly related to foetal state3.
This similarity at the earliest stage is remarkably shown in the course of the arteries which become greatly altered, as fotus advances in life and assumes the widely different course and number which characterize full-grown fish and mammals. How wonderful that in egg, in water or air, or in womb of mother, artery4 should run in same course.
Light can be thrown on this by our theory. The structure of each organism is chiefly adapted to the sustension of its life, when full-grown, when it has to feed itself and propagate5. The structure of a kitten is quite in secondary degree adapted to its habits, whilst fed by its mother's milk and prey. Hence variation in the structure of the full-grown species will chiefly determine the preservation of a
1 See Origin, Ed. i. p. 439, vi. p. 605.
2 In the Origin, Ed. i. p. 440, vi. p. 606, the author argues that the "loop-like course of the arteries" in the vertebrate embryo has no direct relation to the conditions of existence.
3 The following passages are written
across the page:—" They pass through the same phases, but some,
generally called the higher groups, are further metamorphosed.
? Degradation and complication? no tendency to perfection.
? Justly argued against Lamarck? "
4 An almost identical passage occurs in the Origin, Ed. i. p. 440, vi. p. 606.
5 The following: "Deaths of brothers (when) old by same peculiar disease "which is written between the lines seems to have been a memorandum which is expanded a few lines lower. I believe the case of the brothers came from Dr R. W. Darwin.
EMBRYOLOGY 43
species now become ill-suited to its habitat, or rather with a better place opened to it in the economy of Nature. It would not matter to the full-grown cat whether in its young state it was more or less eminently feline, so that it become so when full-grown. No doubt most variation, (not depending on habits of life of individual) depends on early changel and we must suspect that at whatever time of life the alteration of fotus is effected, it tends to appear at same period. When we (see) a tendency to particular disease in old age transmitted by the male, we know some effect is produced during conception, on the simple cell of ovule, which will not produce its effect till half a century afterwards and that effect is not visible2. So we see in grey-hound, bull-dog, in race-horse and carthorse, which have been selected for their form in full-life, there is much less (?) difference in the few first days after birth3, than when full-grown: so in cattle, we see it clearly in cases of cattle, which differ obviously in shape and length of horns. If man were during 10,000 years to be able to select, far more diverse animals from horse or cow, I should expect there would be far less differences in the very young and foetal state: and this, I think, throws light on above marvellous fact. In larvae, which have long life selection, perhaps, does much,—in the pupa not so much4. There is no
1 See the discussion to this effect in the Origin, Ed. i. pp. 443-4, vi. p. 610. The author there makes the distinction between a cause affecting the germ-cell and the reaction occurring at a late period of life.
2 Possibly the sentence was meant to end "is not visible till then."
3 See Origin, Ed. i. pp. 444-5, vi. p. 611. The query appended to much less is justified, since measurement was necessary to prove that the greyhound and bulldog puppies had not nearly acquired "their full amount of proportional difference."
4 (The following discussion, from the back of the page, is in large measure the same as the text.) I think light can be thrown on these facts. From the following peculiarities being hereditary, [we know that some change in the germinal vesicle is effected, which will only betray itself years after] diseases —man, goitre, gout, baldness, fatness, size, [longevity (illegible) time of
44 EMBRYOLOGY
object gained in varying form &c. of foetus (beyond certain adaptations to mother's womb) and therefore selection will not further act on it, than in giving to its changing tissues a tendency to certain parts afterwards to assume certain forms.
Thus there is no power to change the course of
reproduction, shape of horns, case of old brothers dying of same disease]. And we know that the germinal vesicle must have been affected, though no effect is apparent or can be apparent till years afterwards,—no more apparent than when these peculiarities appear by the exposure of the full-grown individual. (That is, "the young individual is as apparently free from the hereditary changes which will appear later, as the young is actually free from the changes produced by exposure to certain conditions in adult life.") So that when we see a variety in cattle, even if the variety be due to act of reproduction, we cannot feel sure at what period this change became apparent. It may have been effected during early age of free life (or) foetal existence, as monsters show. Prom arguments before used, and crossing, we may generally suspect in germ; but I repeat it does not follow, that the change should be apparent till life fully developed; any more than fatness depending on heredity should be apparent during early childhood, still less during fotal existence. In case of horns of cattle, which when inherited must depend on germinal vesicle, obviously no effect till cattle full-grown. Practically it would appear that the [hereditary] peculiarities characterising our domestic races, therefore resulting from vesicle, do not appear with their full characters in very early states; thus though two breeds of cows have calves different they are not so different,—grey-hound and bull-dog. And this is what is (to) be expected, for man is indifferent to characters of young animals and hence would select those full-grown animals which possessed the desirable characteristics. So that from mere chance we might expect that some of the characters would be such only as became fully apparent in mature life. Furthermore we may suspect it to be a law, that at whatever time a new character appears, whether from vesicle, or effects of external conditions, it would appear at corresponding time (see Origin, Ed. i. p. 444). Thus diseases appearing in old age produce children with d°.,—early maturity,—longevity,—old men, brothers, of same disease-young children of d°. I said men do not select for quality of young,—calf with big bullocks. Silk-worms, peculiarities which, appear in caterpillar state or cocoon state, are transmitted to corresponding states. The effect of this would be that if some peculiarity was born in a young animal, but never exercised, it might be inherited in young animal; but if exercised that part of structure would be increased and would be inherited in corresponding time of life after such training.
I have said that man selects in full-life, so would it be in Nature. In struggle of existence, it matters nothing to a feline animal, whether kitten eminently feline, as long as it sucks. Therefore natural selection would act equally well on character which was fully (developed) only in full age. Selection could tend to alter no character in foetus, (except relation to mother) it would alter less in young state (putting on one side larva condition) but alter every part in full-grown condition. Look to a foetus and its parent, and again after ages fotus and its (i.e. the above mentioned parents) descendant; the parent more variable (?) than fotus, which explains all.
RUDIMENTARY ORGANS 45
the arteries, as long as they nourish the foetus; it is the selection of slight changes which supervene at any time during (illegible) of life.
The less differences of foetus,—this has obvious meaning on this view: otherwise how strange that a [monkey] horse, a man, a bat should at one time of life have arteries, running in a manner, which is only intelligibly useful in a fish! The natural system being on theory genealogical, we can at once see, why foetus, retaining traces of the ancestral form, is of the highest value in classification.
§ ix. (ABORTIVE ORGANS.)
There is another grand class of facts relating to what are called abortive organs. These consist of organs which the same reasoning power that shows us how beautifully these organs in some cases are adapted to certain end, declares in other cases are absolutely useless. Thus teeth in Rhinoceros1, whale, narwhal,—bone on tibia, muscles which do not move,—little bone of wing of Apteryx,—bone representing extremities in some snake,—little wings within (?) soldered cover of beetles,—men and bulls, mammae: filaments without anthers in plants, mere scales representing petals in others, in feather-hyacinth whole flower. Almost infinitely numerous. No one can reflect on these without astonishment, can anything be clearer than that wings are to fly and teeth (to bite), and yet we find these organs perfect in every detail in situations where they cannot possibly be of their normal use2.
The term abortive organ has been thus applied
1 Some of these examples occur in Origin, Ed. i. pp. 450-51, vi. PP. 619-20.
2 The two following sentences are written, one down the margin, the other across the page. "Abortive organs eminently useful in classification. Embryonic state of organs. Rudiments of organs."
46 RUDIMENTARY ORGANS
to above structure (as invariable as all other parts1) from their absolute similarity to monstrous cases, where from accident, certain organs are not developed; as infant without arms or fingers with mere stump representing them: teeth represented by mere points of ossification: headless children with mere button,—viscera represented by small amorphous masses, &c.,—the tail by mere stump,— a solid horn by minute hanging one2. There is a tendency in all these cases, when life is preserved, for such structures to become hereditary. We see it in tailless dogs and cats. In plants we see this strikingly,—in Thyme, in Linumflavum,—stamen in Geranium pyrenaicum?. Nectaries abort into petals in Columbine (Aquilegia), produced from some accident and then become hereditary, in some cases only when propagated by buds, in other cases by seed. These cases have been produced suddenly by accident in early growth, but it is part of law of growth that when any organ is not used it tends to diminish (duck's wing4?) muscles of dog's ears, (and of) rabbits, muscles wither, arteries grow up. When eye born defective, optic nerve (Tuco Tuco) is atrophied. As every part whether useful or not (diseases, double flowers) tends to be transmitted to offspring, the origin of abortive organs whether produced at the birth or slowly acquired is easily understood in domestic races of organisms: [a struggle between the atrophy and hereditariness. Abortive organs in domestic races.] There will always be a struggle between atrophy of an organ rendered useless, and
1 I imagine the meaning to be that abortive organs are specific characters in contrast to monstrosities.
2 Minute hanging horns are mentioned in the Origin, Ed. i. p. 454, vi. p. 625, as occurring in hornless breeds of cattle.
3 Linum flavum is dimorphic: thyme gynodisecious. It is not clear what point is referred to under Geranium pyrenaicum.
4 The author's work on duck's wings &c. is in Var. under Dom., Ed. 2, i. p. 299.
RUDIMENTARY ORGANS 47
hereditariness1. Because we can understand the origin of abortive organs in certain cases, it would be wrong to conclude absolutely that all must have had same origin, but the strongest analogy is in favour of it. And we can by our theory, for during infinite changes some organ, we might have anticipated, would have become useless. )We can) readily explain the fact, so astounding on any other view, namely that organs possibly useless have been formed often with the same exquisite care as when of vital importance.
Our theory, I may remark would permit an organ (to) become abortive with respect to its primary use, to be turned to any other purpose, (as the buds in a cauliflower) thus we can see no difficulty in bones of male marsupials being used as fulcrum of muscles, or style of marygold2,—indeed in one point of view, the heads of [vertebrated] animal may be said to be abortive vertebrae turned into other use: legs of some Crustacea abortive jaws, &c., &c. De Candolle's analogy of table covered with dishes3.
(The following passage was possibly intended to be inserted here.) Degradation and complication see Lamarck: no tendency to perfection: if room, [even] high organism would have greater power in beating lower one, thought (?) to be selected for a degraded end.
1 The words vis medicatrix are inserted after "useless," apparently as a memorandum.
2 In the male florets of certain Composites the style functions merely as a piston for forcing out the pollen.
3 (On the back of the page is the following.) If abortive organs are a trace preserved by hereditary tendency, of organ in ancestor of use, we can at once see why important in natural classification, also why more plain in young animal because, as in last section, the selection has altered the old animal most. I repeat, these wondrous facts, of parts created for no use in past and present time, all can by my theory receive simple explanation; or they receive none and we must be content with some such empty metaphor, as that of De Candolle, who compares creation to a well covered table, and says abortive organs may be compared to the dishes (some should be empty) placed symmetrically!
48 RECAPITULATION
§ x. RECAPITULATION AND CONCLUSION.
Let us recapitulate the whole (?) (of) these latter sections by taking case of the three species of Rhinoceros, which inhabit Java, Sumatra, and mainland of Malacca or India. We find these three close neighbours, occupants of distinct but neighbouring districts, as a group having a different aspect from the Rhinoceros of Africa, though some of these latter inhabit very similar countries, but others most diverse stations. We find them intimately related [scarcely (?) differences more than some breeds of cattle] in structure to the Rhinoceros, which for immense periods have inhabited this one, out of three main zoological divisions of the world. Yet some of these ancient animals were fitted to very different stations: we find all three (illegible) of the generic character of the Rhinoceros, which form a [piece of net]1 set of links in the broken chain representing the Pachydermata, as the chain likewise forms a portion in other and longer chains. We see this wonderfully in dissecting the coarse leg of all three and finding nearly the same bones as in bat's wings or man's hand, but we see the clear mark in solid tibia of the fusion into it of the fibula. In all three we find their heads composed of three altered vertebrae, short neck, same bones as giraffe. In the upper jaws of all three we find small teeth like rabbit's. In dissecting them in fotal state we find at a not very early stage their form exactly alike the most different animals, and even with arteries running as in a fish: and this similarity holds when the young one is produced in womb, pond, egg or spawn. Now these three undoubted species scarcely differ more than breeds of cattle,
1 The author doubtless meant that the complex relationships between organisms can be roughly represented by a net in which the knots stand for species.
RECAPITULATION 49
are probably subject to many the same contagious diseases; if domesticated these forms would vary, and they might possibly breed together, and fuse into something1 different (from) their aboriginal forms; might be selected to serve different ends.
Now the Creationist believes these three Rhinoceroses were created2 with their deceptive appearance of true, not (illegible) relationship; as well can I believe the planets revolve in their present courses not from one law of gravity but from distinct volition of Creator.
If real species, sterile one with another, differently adapted, now inhabiting different countries, with different structures and instincts, are admitted to have common descent, we can only legitimately stop where our facts stop. Look how far in some case a chain of species will lead us. (This probably refers to the Crustacea, where the two ends of the series have "hardly a character in common." Origin, Ed. i. p. 419.) May we not jump (considering how much extermination, and how imperfect geological records) from one sub-genus to another sub-genus. Can genera restrain us; many of the same arguments, which made us give up species, inexorably demand genera and families and orders to fall, and classes tottering. We ought to stop only when clear unity of type, independent of use and adaptation, ceases.
Be it remembered no naturalist pretends to give test from external characters of species; in many genera the distinction is quite arbitrary3. But there remains one other way of comparing species
1 Between the lines occurs:—"one (?) form be lost."
2 The original sentence is here broken up by the insertion of:—"out of the dust of Java, Sumatra, these (?) allied to past and present age and (illegible), with the stamp of inutility in some of their organs and conversion mothers."
Between the lines occur the words:—" Species vary according to same general laws as varieties; they cross according to same laws."
50 CONCLUSION
with races; it is to compare the effects of crossing them. Would it not be wonderful, if the union of two organisms, produced by two separate acts of Creation, blended their characters together when crossed according to the same rules, as two races which have undoubtedly descended from same parent stock; yet this can be shown to be the case. For sterility, though a usual (?), is not an invariable concomitant, it varies much in degree and has been shown to be probably dependent on causes closely analogous with those which make domesticated organisms sterile. Independent of sterility there is no difference between mongrels and hybrids, as can be shown in a long series of facts. It is strikingly seen in cases of instincts, when the minds of the two species or races become blended together1. In both cases if the half-breed be crossed with either parent for a few generations, all traces of the one parent form is lost (as Kolreuter in two tobacco species almost sterile together), so that the Creationist in the case of a species, must believe that one act of creation is absorbed into another!
[-page break-]
[-page break-]
CONCLUSION.
Such are my reasons for believing that specific forms are not immutable. The affinity of different groups, the unity of types of structure, the representative forms through which foetus passes, the metamorphosis of organs, the abortion of others cease to be metaphorical expressions and become intelligible facts. We no longer look (an) on animal as a savage does at a ship2, or other great work of art, as a thing wholly beyond comprehension, but we
1 "A cross with a bull-dog has affected for many generations the courage and obstinacy of greyhounds," Origin, Ed. i. p. 214, vi. p. 327.
2 The simile of the savage and the ship occurs in the Origin, Ed. i. p. 485, vi. p. 665.
CONCLUSION 51
feel far more interest in examining it. How interesting is every instinct, when we speculate on their origin as an hereditary or congenital habit or produced by the selection of individuals differing slightly from their parents. We must look at every complicated mechanism and instinct, as the summary of a long history, (as the summing up) of1 useful contrivances, much Hike a work of art. How interesting does the distribution of all animals become, as throwing light on ancient geography. [We see some seas bridged over.] Geology loses in its glory from the imperfection of its archives2, but how does it gain in the immensity of the periods of its formations and of the gaps separating these formations. There is much grandeur in looking at the existing animals either as the lineal descendants of the forms buried under thousand feet of matter, or as the coheirs of some still more ancient ancestor. It accords with what we know of the law impressed on matter by the Creator, that the creation and extinction of forms, like the birth and death of individuals should be the effect of secondary [laws] means3. It is derogatory that the Creator of countless systems of worlds should have created each of the myriads of creeping parasites and [slimy] worms which have swarmed each day of life on land and water (on) [this] one globe. We cease being astonished, however much we may deplore, that a group of animals should have been directly created to lay their eggs in bowels and flesh of other,—that some organisms should delight in cruelty,—that animals should be led away by false instincts,—that annually there should be an
1 In the Origin, Ed. i. p. 486, vi. p. 665, the author speaks of the "summing up of many contrivances": I have therefore introduced the above words which make the passage clearer. In the Origin the comparison is with "a great mechanical invention,"-not with a work of art. eee a similar passage in the Origin, Ed. i. p. 487, vi. p. 667.
2 See the Origin, Ed. i. p. 488, vi. p. 668.
52 CONCLUSION
incalculable waste of eggs and pollen. From death, famine, rapine, and the concealed war of nature we can see that the highest good, which we can conceive, the creation of the higher animals has directly come. Doubtless it at first transcends our humble powers, to conceive laws capable of creating individual organisms, each characterised by the most exquisite workmanship and widely-extended adaptations. It accords better with [our modesty] the lowness of our faculties to suppose each must require the fiat of a creator, but in the same proportion the existence of such laws should exalt our notion of the power of the omniscient Creator1. There is a simple grandeur in the view of life with its powers of growth, assimilation and reproduction, being originally breathed into matter under one or a few forms, and that whilst this our planet has gone circling on according to fixed laws, and land and water, in a cycle of change, have gone on replacing each other, that from so simple an origin, through the process of gradual selection of infinitesimal changes, endless forms most beautiful and most wonderful have been evolved2.
1 The following discussion, together with some memoranda are on the last page of the ms. "The supposed creative spirit does not create either number or kind which (are) from analogy adapted to site (viz. New Zealand): it does not keep them all permanently adapted to any country,—it works on spots or areas of creation,—it is not persistent for great periods,—it creates forms of same groups in same regions, with no physical similarity,—it creates, on islands or mountain summits, species allied to the neighbouring ones, and not allied to alpine nature as shown in other mountain summits -even different on different island of similarly constituted archipelago, not created on two points: never mammifers created on small isolated island; nor number of organisms adapted to locality: its power seems influenced or related to the range of other species wholly distinct of the same genus,—it does not equally effect, in amount of difference, all the groups of the same class."
2 This passage is the ancestor of the concluding words in the first edition of the Origin of Species which have remained substantially unchanged throughout subsequent editions, "There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most
CONCLUSION 53
N.B.—There ought somewhere to be a discussion from Lyell to show that external conditions do vary, or a note to Lyell's works (work?).
Besides other difficulties in ii. Part, non-acclimatisation of plants. Difficulty when asked how did white and negro become altered from common intermediate stock: no facts. We do not know that species are immutable, on the contrary. What arguments against this theory, except our not perceiving every step, like the erosion of valleys1.
beautiful and most wonderful have been, and are being, evolved." In the 2nd edition "by the Creator" is introduced after "originally breathed."
1 Compare the Origin, Ed. i. p. 481, vi. p. 659, "The difficulty is the same as that felt by so many geologists, when Lyell first insisted that long lines of inland cliffs had been formed, and great valleys excavated, by the slow action of the coast-waves."