"
I may finally call attention to the probability that the
association of paternal and maternal chromosomes in pairs
and their subsequent separation during the reducing division
... may constitute the physical basis of the Mendelian law
of heredity." [emphasis added]. W Sutton (1902)
Biol. Bull. 4: 24-39:
"
1. The chromosome group of the presynaptic germ-cells
is made up of two equivalent chromosome-series, and that
strong ground exists for the conclusion that one of
these is paternal and the other maternal.
[Fig.1]
2. The process of synapsis
(pseudo-reduction) consists in the union in pairs of the homologous
members (i. e., those that correspond in size) of
the two series. [Fig. 5a]
3. The first post-synaptic
or maturation mitosis is equational
and hence results in no chromosomic
differentiation. [Fig. 5b]
4. The second post-synaptic
division is a reducing division, resulting in the
separation of the chromosomes which have conjugated in
synapsis, and their relegation to different germ-cells [Fig.
9].
5. The chromosomes retain a
morphological individuality throughout the
various cell-divisions."
W. S. Sutton (1903)
The Chromosomes in Heredity Biol. Bull. 4: 231-251 [emphasis
added]