"How the giraffe got
its neck"
"It is interesting to observe the result of habit in the peculiar shape and size of the giraffe (Camelo-pardalis): this animal, the largest of the mammals, is know to live in the interior of Africa in places where the soil is nearly always arid and barren, so that it is obliged [il besoin] to browse on the leaves of trees and to make constant efforts to reach them. From this habit long maintained in all its race, it has resulted that the animal's fore-legs have become longer than its hind legs, and that its neck is lengthened to such a degree that the giraffe, without standing up on its hind legs, attains a height of six metres (nearly 20 feet)." - Jean Baptiste Lamarck (1809) "Zoological Philosophy"
Lamarck's explanation of the giraffe's neck is a classic "adaptationist" story
[sometimes called "Just
So" stories after those of Rudyard Kipling],
a clever and attractive explanation of the adaptive significance
of a character, but onen that is not based on any empirical
data. Observation of giraffes shows that they actually prefer to
browse at shoulder height [below, left]. The extended neck of
giraffes instead appears to be a consequence of sexual selection. Male
giraffes use their neck and head as clubs in agonistic displays
with other males in competition for females [below, right]. Data
demonstrate that those with thicker necks and more massive
skulls and horns are more successful, and are the mates
preferred by females.