
          
        Theodosius
            Dobzhansky (1900 - 1975)
      Феодо́сий
            Григо́рьевич Добржа́нский 
          The Ukrainian-born biologist was an early
      proponent of the study of genetic variation in natural populations
      as a means to understand evolution. After leaving the Soviet
      Union, he went first to Cal Tech in southern California and then
      to Columbia University in New York. he studied chromosomal
      variation in Drosophila pseudoobscura, a wild relative of
      D. melanogaster. His demonstration that frequencies of
      different chromosomal arrangements within species varied according
      to altitude, temperature, and humidity, and regularly cycled over
      the seasons of the year, was the first clear evidence that genetic
      variation was adaptive, as Darwin had predicted. His "Genetics
          and the Origin of Species" (1937; 4th ed., 1970)
      contributed the Genetics component of the Modern Synthesis.
      His aphorism that "Nothing in biology
            makes sense except in the light of evolution"
      is widely quoted.